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  1. Syfert, M.M., Smith, M.J., Coomes, D.A. (2013) The effects of sampling bias and model complexity on the predictive performance of MaxEnt species distribution models. PLoS ONE 8(2): e55158. DOI link
  2. Vanderwel M.C, Coomes,D.A., and Purves, D.W. (2013) Quantifying variation in forest disturbance, and its effects on aboveground biomass dynamics, across the eastern United States. Global Change Biology DOI link
  3. Ruiz-Benito, P., Lines, E.R., Gómez-Aparicio, L., Zavala, M.A., Coomes, D.A. (2013) Patterns and Drivers of Tree Mortality in Iberian Forests: Climatic Effects Are Modified by Competition. PLoS ONE 8(2): e56843. DOI link


  1. Tanentzap, A. J., Mountford, E. P., Cooke, A. S., & Coomes, D.A. (2012). The more stems the merrier: advantages of multi-stemmed architecture for the demography of understorey trees in a temperate broadleaf woodland. Journal of Ecology. DOI link
  2. Tanentzap, A. J., & Coomes, D.A. (2012) Carbon storage in terrestrial ecosystems: Do browsing and grazing herbivores matter? Biological Reviews, 87(1), 72-94.
  3. Price, C. A., Weitz, J. S., Savage, V. M., Stegen, J., Clarke, A., Coomes, D.A., … Swenson, N. G. (2012) Testing the metabolic theory of ecology. Ecology Letters. DOI link
  4. Brodribb, T. J., Pittermann, J., & Coomes, D.A. (2012) Elegance versus speed: Examining the competition between conifer and angiosperm trees. International Journal of Plant Sciences, 173, 673-694. R
  5. Kunstler, G., Lavergne, S., Courbaud, B., Thuiller, W., Vieilledent, G., Zimmermann, N. E., Coomes, D.A. (2012) Competitive interactions between forest trees are driven by species' trait hierarchy, not phylogenetic or functional similarity: Implications for forest community assembly. Ecology Letters, 15(8), 831-840.
  6. Simonson, W. D., Allen, H. D., & Coomes, D.A. (2012) Use of an airborne lidar system to model plant species composition and diversity of Mediterranean oak forests. Conservation Biology, 26, 840-850.
  7. Tanentzap, A. J., Lee, W. G., & Coomes, D.A. (2012) Soil nutrient supply modulates temperature-induction cues in mast-seeding grasses. Ecology, 93, 462-469.
  8. Coomes, D.A., Holdaway, R. J., Kobe, R. K., Lines, E. R., & Allen, R. B. (2012) A general integrative framework for modelling woody biomass production and carbon sequestration rates in forests. Journal of Ecology, 100, 42-64.
  9. Lines, E. R., Zavala, M. A., Purves, D. W., & Coomes, D.A. (2012) Predictable changes in aboveground allometry of trees along gradients of temperature, aridity and competition. Global Ecology and Biogeography. PAGE
  10. Wright, D. M., Tanentzap, A. J., Flores, O., Husheer, S. W., Duncan, R. P., Wiser, S. K., … Coomes, D.A. (2012) Impacts of culling and exclusion of browsers on vegetation recovery across New Zealand forests. Biological Conservation, 153, 64-71.
  11. Echeverría, C., Newton, A., Nahuelhual, L., Coomes, D.A., & Rey-Benayas, J. M. (2012) How landscapes change: Integration of spatial patterns and human processes in temperate landscapes of southern Chile. Applied Geography, 32(2), 822-831.
  12. Jucker, T., & Coomes, D.A. (2012) Comment on "Plant species richness and ecosystem multifunctionality in global drylands". Science, 337(6091), 155.


  1. Hurst, J. M., Allen, R. B., Coomes, D.A., & Duncan, R. P. (2011). Size-specific tree mortality varies with neighbourhood crowding and disturbance in a Montane Nothofagus forest. PLoS One, 6(10), e26670. doi:10.1371/journal.pone.0026670
  2. Echeverría, C., Newton, A., Nahuelhual, L., Coomes, D., & Rey-Benayas, J. M. (2012). How landscapes change: Integration of spatial patterns and human processes in temperate landscapes of southern Chile. Applied Geography, 32(2), 822-831.
  3. Kattge, J., Díaz, S., Lavorel, S., Prentice, I. C., Leadley, P., Bönisch, G., …Coomes, D.A... Wirth, C. (2011). TRY - a global database of plant traits. Global Change Biology, 17(9), 2905-2935.
  4. Hoorens, B., Coomes, D., & Aerts, R. (2010). Neighbour identity hardly affects litter-mixture effects on decomposition rates of New Zealand forest species. Oecologia, 162(2), 479-489.
  5. Rands, M. R. W., Adams, W. M., Bennun, L., Butchart, S. H. M., Clements, A., Coomes, D.… Vira, B. (2011). Culture and Biodiversity Losses Linked Response. Science, 331(6013), 31.
  6. Rands, M. R., Adams, W. M., Bennun, L., Butchart, S. H., Clements, A., Coomes, D., … Vira, B. (2010). Biodiversity conservation: challenges beyond 2010. Science, 329(5997), 1298-1303. DOI link
  7. Tanentzap, A. J., & Coomes, D.A. (2011). Carbon storage in terrestrial ecosystems: do browsing and grazing herbivores matter?. Biological Reviews. DOI link
  8. Coomes, D.A., Lines, E. R., & Allen, R. B. (2011). Moving on from Metabolic Scaling Theory: hierarchical models of tree growth and asymmetric competition for light. Journal of Ecology, 99(3), 748-756. DOI link
  9. Flores, O., & Coomes, D.A. (2011). Estimating the wood density of species for carbon stock assessments. Methods in Ecology and Evolution, 2(2), 214-220. DOI link
  10. Kunstler, G., Albert, C. H., Courbaud, B., Lavergne, S., Thuiller, W., Vieilledent, G. … Coomes, D.A. (2011). Effects of competition on tree radial-growth vary in importance but not in intensity along climatic gradients. Journal of Ecology, 99(1), 300-312. DOI link
  11. Bee, J. N., Tanentzap, A. J., Lee, W. G., Lavers, R. B., Mark, A. F., Mills, J. A., ... Coomes, D.A. (2011). Influence of foliar traits on forage selection by introduced red deer in New Zealand. Basic and Applied Ecology, 12(1), 56-63.
  12. Tanentzap, A. J., Bazely, D. R., Koh, S., Timciska, M., Haggith, E. G., Carleton, T. J., Coomes, D.A. (2011). Seeing the forest for the deer: Do reductions in deer-disturbance lead to forest recovery?. Biological Conservation, 144(1), 376-382.
  13. Tanentzap, A. J., Lee, W. G., Dugdale, J. S., Patrick, B. P., Fenner, M., Walker, S., Coomes, D.A. (2011). Differential responses of vertebrate and invertebrate herbivores to traits of New Zealand subalpine shrubs. Ecology, 92(4), 994-999.
  14. Holdaway, R. J., Richardson, S. J., Dickie, I. A., Peltzer, D.A., & Coomes, D.A. (2011). Species- and community-level patterns in fine root traits along a 120000-year soil chronosequence in temperate rain forest. Journal of Ecology, 99(4), 954-963.1466. DOI link
  15. Spence, L. A., Dickie, I. A., & Coomes, D.A. (2011). Arbuscular mycorrhizal inoculum potential: a mechanism promoting positive diversity-invasibility relationships in mountain beech forests in New Zealand?. Mycorrhiza, 21(4), 309-314. DOI link


  1. Bystriakova, N., Bader, M. and Coomes, D.A. (2011) Long-term tree fern dynamics linked to disturbance and shade tolerance. Journal of Vegetation Science 22, 72–84. DOI link
  2. Bystriakova, N., Schneider, H., & Coomes, D. (2010). Evolution of the climatic niche in scaly tree ferns (Cyatheaceae, Polypodiopsida). Botanical Journal of the Linnean Society, 165(1), 1-19. DOI link
  3. Bee, J.N., Tanentzap, A.T, Lee, W.G., Lavers, R.B., Mark, A.F., Mills, J.A., Coomes, D.A , Influence of foliar traits on forage selection by introduced red deer in New Zealand. Basic and Applied Ecology, DOI link.
  4. Rands, M.R.W., Adams, W.M., Bennun, L., Butchart, S.H.M., Clements, A., Coomes, D.A., Entwistle, A., Hodge, I., Kapos, V., Scharlemann, J.P.W., Sutherland, W.J., Vira, B. (2010) Biodiversity Conservation: Challenges Beyond 2010. Science 329: 1298- 1303. DOI link
  5. Lafortezza, R., Coomes, D.A., Kapos, V., & Ewers, R.M. (2010) Assessing the impacts of fragmentation on plant communities in New Zealand: scaling from survey plots to landscapes. Global Ecology and Biogeography 19, 741–75. DOI link
  6. Wright, D.M., Jordan, G.J., Lee, W.G., Duncan, R.P., Forsyth, D.M., & Coomes, D.A. (2010) Do leaves of plants on phosphorus-impoverished soils contain high concentrations of phenolic defence compounds? Functional Ecology 24, 52-61. DOI link
  7. Forsyth, D.M., Wilmshurst, J.M., Allen, R.B., & Coomes, D.A. (2010) Impacts of introduced deer and extinct moa on New Zealand ecosystems. New Zealand Journal of Ecology 34, 48–65. click
  8. Russo S.E., Jenkins K.L., Wiser S.K., Uriarte M., Duncan R.P., Coomes D.A. (2010) Interspecific relationships among growth, mortality and xylem traits of woody species from New Zealand. Functional Ecology 24: 253-262.
  9. Flores, O., Coomes, D.A.(2010) Estimating the wood density of species for carbon stock assessments. Methods in Ecology and Evolution DOI link
  10. Gaxiola A., McNeill S.M., Coomes, D.A. (2010) What drives retrogressive succession? Plant strategies to tolerate infertile and poorly drained soils. Functional Ecology 24:714–722. DOI link
  11. Lines E.R., Coomes D.A., Purves D.W. (2010) Influences of Forest Structure, Climate and Species Composition on Tree Mortality across the Eastern US. PLoS ONE 5(10): e13212. DOI link
  12. Spence, L.A., Dickie, I.A., Coomes, D.A.(2010) Arbuscular mycorrhizal inoculum potential: a mechanism promoting positive diversity–invasibility relationships in mountain beech forests in New Zealand? DOI link
  13. Bee, J.N., Wright, D.M., Tanentzap, A.J., Lee, W.G., Lavers, R.B., Mark, A.F., Mills, J.A., & Coomes, D.A. (2010) Spatio-temporal feeding selection of red deer in a mountainous landscape. Austral Ecology, 35, 752–764. DOI link
  14. Spence, L.A., Ross, J.V., Wiser, S.K., Allen, R.B. & Coomes, D.A. (2010) Disturbance affects short-term facilitation, but not long-term saturation, of exotic plant invasion in New Zealand forest. Proceeding Royal Society - Biology DOI link
  15. Tanentzap, A.J., Lee, W.G., Dugdale, J.S., Patrick, B.P., Fenner, M., Walker, S., & Coomes, D.A. (2010) Differential responses of vertebrate and invertebrate herbivores to traits of New Zealand subalpine shrubs. Ecology, in press. DOI link
  16. Tanentzap, A.J., Bazely, D.R., Koh, S., Timciska, M., Haggith, E.G., Carleton, T.J., & Coomes, D.A. (2010)Seeing the forest for the deer: Do reductions in deer-disturbance lead to forest recovery? Biological Conservation, in press. DOI link


  1. Coomes, D.A. & Sack, L. (2009) Response to comment on Coomes et al. ‘Scaling of xylem vessels and veins within the leaves of oak species’. Biology Letters, 5, 381-382.
  2. Hoorens, B, Coomes, D.A., Aerts, R. (2009) Neighbour identity hardly affects litter mixture effects on decomposition rates of New Zealand forest species. Oecologia 162, 479-89.
  3. Kunstler, G., Coomes, D.A., Canham, C.D. (2009) Size-dependence of growth and mortality influence the shade tolerance of trees in a lowland temperate rain forest. Journal of Ecology. 97, 685 - 695. DOI link
  4. Coomes, D.A., Kunstler, G., Canham, C.D., & Wright, E. (2009) A greater range of shade-tolerance niches in nutrient-rich forests: an explanation for positive richness–productivity relationships? Journal of Ecology. 97, 705-717. external link: DOI link pdf (password needed)
  5. Chave, J., Coomes,D.A, Jansen, S., Lewis, S., Swenson, N., Zanne, A. (2009) Towards a worldwide wood economics spectrum. Ecology Letters, 12, 351-366 DOI link pdf
  6. Ewers, R.M., Kapos, V., & Coomes, D.A., Lafortezza, R., & Didham, R.K. (2009) Mapping community change in modified landscapes. Biological Conservation, 142, 2872-2880. DOI link
  7. Tanentzap, A.J., Burrows, L.E., Lee, W.G., Nugent, G., Maxwell, J.M., & Coomes, D.A. (2009) Landscape-level vegetation recovery from herbivory: progress after four decades of invasive red deer control. Journal of Applied Ecology 46, 1064–1072. DOI link
  8. Tanentzap, A.J., Bee, J.N., Lee, W.G., Lavers, R.B., Mills, J.A., Mark. A.F., & Coomes D.A. (2009) The reliability of palatability estimates obtained from rumen contents analysis and a field-based index of diet selection. Journal of Zoology. 278, 243–248. DOI link pdf (password needed)
  9. Bee, J.N, Tanentzap, A.J., Lee, W.G., Lavers, R.B., Mark, A.F., Mills, J.A., Coomes, D.A. (2009) The benefits of being in a bad neighbourhood: plant community composition influences red deer foraging decisions. Oikos, 118, 18-24. DOI link
  10. Echeverría, E., Coomes, D.A., Hall M. and Newton, A.C. (2008) Spatially explicit models to analyze forest loss and fragmentation between 1976 and 2020 in southern Chile. Ecological Modelling DOI link pdf (password needed)


  1. Coomes, D.A., Heathcote, S., Godfrey, E.R., Shepherd, J.J., and Sack, L. (2008) Scaling of xylem vessels and veins within the leaves of oak species. Biology Letters 4, 302-306 DOI link pdf (password needed)
  2. Weedon, J., Cornwell, W., Cornelissen, H., Zanne, A., Wirth, C., Coomes, D.A. (2008). Global meta-analysis of wood decomposition rates: a role for trait variation among tree species? Ecology Letters 12, 45-56. DOI link
  3. Janssen, T., Bystriakova, N., Rakotondrainibe, F., Coomes, D.A.,, Labat, N.L., Schneider, H. (2008) Neoendemism in Madagascan scaly tree ferns results from recent, coincident diversification bursts. Evolution 62: 1876-89. DOI link
  4. Grubb,P.J., Jackson, R.V., Barberis, I.M., Bee, J.N., Coomes, D.A., Dominy, N.J., De La Fuente, M.A.S. Lucas, P.W., Metcalfe, D.J., Svenning, J.C., Turner, I.M. & Vargas O. (2008) Monocot leaves are eaten less than dicot leaves in tropical lowland rain forests: correlations with toughness and leaf presentation. Annals of Botany 101, 1379-1389 DOI link pdf (password needed)
  5. Gaxiola, A., Burrows, L.E. & Coomes D.A. (2008) Tree-fern trunks facilitate seedling regeneration in a productive lowland temperate rain forest. Oecologia, 155, 325–335. DOI link pdf (password needed)
  6. Holdaway, R.J., Allen, R.B., Clinton, P.W., Davis, M.R. & Coomes,D.A.(2008) Intraspecific changes in forest canopy allometries during self-thinning. Functional Ecology DOI link pdf (password needed)


  1. Turnbull, L.A, Coomes, D.A., Purves, D.W., & Rees, M. (2007) How spatial structure alters population and community dynamics in a natural plant community. Journal of Ecology 95, 79–89. DOI link
  2. Kunstler, G., Allen, R.B., Coomes, D.A., Canham, C.D., & Wright, E.F. (2007) Long-term harvesting of podocarps. Landcare Research Contract Report LC0607/188
  3. Coomes, D.A. & Allen, R.B. (2007) Effects of size, competition and altitude on tree growth. Journal of Ecology 95, 1084–1097. DOI link pdf (password needed)
  4. Coomes, D.A & Allen, R.B. (2007) Mortality and tree-size distributions in natural mixed-age forests. Journal of Ecology, 95, 27-40. DOI link
  5. Coomes, D.A., Jenkins, K.L., & Cole, L.E.S. (2007) Scaling of tree vascular transport systems along gradients of nutrient supply and altitude. Biology Letters, 3, 86-89. DOI link
  6. Bee, J. N., Kunstler G. & Coomes, D.A. (2007) Resistance and resilience of New Zealand tree species to browsing. Journal of Ecology 95, 1014–1026. DOI link pdf (password needed)
  7. Echeverría, C., Newton, A.C., Lara, A., Benayas, J.M.R., & Coomes, D.A. (2007) Impacts of forest fragmentation on species composition and forest structure in the temperate landscape of southern Chile. Global Ecology and Biogeography, 16, 426-439. DOI link


  1. Coomes, D.A. (2006) Challenges to the generality of WBE scaling theory. Trends In Ecology & Evolution, 21, 593-596. DOI link
  2. Russo, S., Wiser, S.W., & Coomes, D.A. (2007) Growth-size scaling relationships of woody plant species differ from predictions of the Metabolic Theory of Ecology. Ecology Letters, 10, 889–901. DOI link
  3. Parfitt, R.L., Ross, D.J., & Coomes, D.A., Richardson, S.J., Smale, M.C., & Dahlgren, R.A. (2005) N and P in New Zealand soil chronosequences and relationships with foliar N and P. Biogeochemistry, 75, 305-328. click
  4. Echeverría, C., Cayuela, L., Manson, R., Coomes, D.A., Lara, A. , Rey-Benayas, J.M. & Newton A.C. (2007) Spatial and temporal patterns of forest loss and fragmentation in Mexico and Chile. In: Biodiversity Loss and Conservation in Fragmented Forest Landscapes: The Forests of Montane Mexico and Temperate South America (ed A.C. Newton), pp 14-41. CABI, Wallingford, Oxford.
  5. Echeverría, C., Coomes, D.A., Salas, J., Rey-Benayas, J.M., Lara, A., & Newton, A. (2006) Rapid deforestation and fragmentation of Chilean Temperate Forests. Biological Conservation, 130, 481-494. DOI link
  6. Coomes, D.A., Mark, A.F., & Bee, J. (2006) Animal Control and Ecosystem Recovery. In: Biological Invasions in New Zealand (eds R.B. Allen & W.G. Lee), pp. 339-354, Springer Verlag.
  7. Husheer, S.W., Robertson, A.W., Coomes, D.A., & Frampton, C.M. (2006) Herbivory and plant competition reduce mountain beech seedling growth and establishment in New Zealand. Plant Ecology, 183, 245-256. DOI link

2005 and earlier

  1. Coomes, D.A., Allen, R.B., Bentley, W.A., Burrows, L.E., Canham, C.D., Fagan, L., Forsyth, D.M., Gaxiola-Alcantar, A., Parfitt, R.L., Ruscoe, W.A., Wardle, D.A., Wilson, D.J., & Wright, E.F. (2005) The hare, the tortoise and the crocodile: the ecology of angiosperm dominance, conifer persistence and fern filtering. Journal of Ecology, 93, 918-935. click
  2. Grubb, P.J., Metcalfe, D.J. & Coomes, D.A. (2005) Comment on "A brief history of seed size". Science, 310, 783A-783A. click
  3. Tanner, E.V.J., Teo, V.K., Coomes, D.A. & Midgley, J.J. (2005) Pair-wise competition-trials amongst seedlings of ten dipterocarp species; the role of initial height, growth rate and leaf attributes. Journal of Tropical Ecology, 21, 317-328. click
  4. Platt, K.H., Allen, R.B., Coomes, D.A., & Wiser, S.K. (2004) Mountain beech seedling responses to removal of below-ground competition and fertiliser addition. New Zealand Journal of Ecology, 28, 289-293. click
  5. Turnbull, L.A., Coomes, D.A., Hector, A., & Rees, M. (2004) Seed mass and the competition/colonization trade-off: competitive interactions and spatial patterns in a guild of annual plants. Journal of Ecology, 92, 97-109. click
  6. Husheer, S.W., Coomes, D.A., & Robertson, A.W. (2003) Long-term influences of introduced deer on the composition and structure of New Zealand Nothofagus forests. Forest Ecology and Management, 181, 99-117. click
  7. Coomes, D.A., Allen, R.B., Scott, N.A., Goulding, C., Beets, P. (2003) Designing systems to monitor carbon stocks in forests and shrublands. Forest Ecology and Management, 164, 89-108. click
  8. Bellingham, P.J. & Coomes, D.A. (2003) Grazing and community structure as determinants of invasion success by Scotch broom in a New Zealand montane shrubland. Diversity and Distributions, 9, 19-28. click
  9. Coomes, D.A., Duncan, R.P., Allen, R.B., & Truscott, J. (2003) Disturbances prevent stem size-density distributions in natural forests from following scaling relationships. Ecology Letters, 6, 980-989. click
  10. Coomes, D.A., Allen, R.B., Forsyth, D.M. & Lee, W.G. (2003) Factors preventing the recovery of New Zealand forests following control of invasive deer. Conservation Biology, 17, 450-459. click
  11. Forsyth, D.M., Coomes, D.A., & Nugent, G. (2003) Framework for assessing the susceptibility of management areas to deer impacts. Science for Conservation Series, Issue 213. 39 pages. Department of Conservation, Wellington, New Zealand.
  12. Coomes, D.A., & Grubb, P.J. (2003) Colonization, tolerance, competition and seed-size variation within functional groups. Trends in Ecology & Evolution, 18, 283-291. click
  13. Wardle, D.A., Hornberg, G., Zackrisson, O., Kalela-Brundin, M., & Coomes, D.A. (2003) Long-term effects of wildfire on ecosystem properties across an island area gradient. Science, 300, 972-975. click
  14. Coomes, D.A., Rees, M., Grubb, P.J., & Turnbull, L. (2002) Are differences in seed mass among species important in structuring plant communities? Evidence from analyses of spatial and temporal variation in dune-annual populations. Oikos, 96, 421-432. click
  15. Coomes, D.A., Rees, M., Turnbull, L., & Ratcliffe, S. (2002) On the mechanisms of coexistence among annual-plant species, using neighbourhood techniques and simulation models. Plant Ecology, 163, 23-38. click
  16. Coomes, D.A. & Grubb, P.J. (2000) Impacts of root competition in forests and woodlands: A theoretical framework and review of experiments. Ecological Monographs, 70, 171-207. click
  17. Coomes, D.A., Rees, M., & Turnbull, L. (1999) Identifying aggregation and association in fully mapped spatial data. Ecology, 80, 554-565. click
  18. Kollmann, J., Coomes, D.A., & White, S.M. (1998) Consistencies in post-dispersal seed predation of temperate fleshy-fruited species among seasons, years and sites. Functional Ecology, 12, 683-690. click
  19. Coomes, D.A. & Grubb, P.J. (1998) A comparison of 12 tree species of Amazonian caatinga using growth rates in gaps and understorey, and allometric relationships. Functional Ecology, 12, 426-435. click
  20. Coomes, D.A. & Grubb, P.J. (1998) Responses of juvenile trees to above- and belowground competition in nutrient-starved Amazonian rain forest. Ecology, 79, 768-782. click
  21. Bonis, A., Grubb, P.J., & Coomes, D.A. (1997) Requirements of gap-demanding species in chalk grassland: reduction of root competition versus nutrient-enrichment by animals. Journal of Ecology, 85, 625-633. click
  22. Coomes, D.A. (1997) Nutrient status of Amazonian caatinga forests in a seasonally dry area: nutrient fluxes in litter fall and analyses of soils. Canadian Journal of Forest Research, 27, 831-839. click
  23. Grubb, P.J. & Coomes, D.A. (1997) Seed mass and nutrient content in nutrient-starved tropical rainforest in Venezuela. Seed Science Research, 7, 269-280. click
  24. Coomes, D.A. & Grubb, P.J. (1996) Amazonian caatinga and related communities at La Esmeralda, Venezuela: Forest structure, physiognomy and floristics, and control by soil factors. Vegetatio, 122, 167-191. click


  1. Wiser, S., Allen, R., Peltzer, D., Benecke, U, Baker, G., & Coomes, D.A (2003) Growth and survival of residual trees after harvesting. Landcare Research Contract Report LC0203/125, prepared for Ministry of Agriculture and Forestry, New Zealand.
  2. Svavarsdóttir, K., Allen, R.B., Burrows, L., Coomes, D.A., Wiser, S., Smale M., & Benecke, U (1999) Silvacultural research in selected forest types: a podocarp-tawa forest in Taranaki. Landcare Research Contract Report LC9899/119, prepared for Ministry of Agriculture and Forestry, New Zealand.
  3. Allen, R, Burley, J., Coomes, D.A, Beets, P., Goulding, C. (1999) Data handling and database design: A national system for monitoring changes in carbon stored in indigenous forests and scrub: data models and linkages to software for South Island transect. Landcare Research Contract Report JNT9899/125, prepared for Ministry for the Environment, New Zealand.